From evolutionary point of view, retention of the female gametophyte with developing young embryo on the parent sporophyte for some time, is first observed in
Liverworts
Mosses
Pteridophytes
Gymnosperms
In Pteridophyte, megaspore is retained for some times in female gametophyte, however the permanent retention is required for seed formation in Gymnosperms. That's why Pteridophytes exhibit precursor to seed habit only.
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