Pinus seed cannot germinate and established without fungal association. This is because :
its embryo is immature.
it has obligate association with mycorrhizae.
it has very hard seed coat.
its seeds contain inhibitors that present germination.
Fungus associated with roots of Pinus increases minerals & water absorption for the plant by increasing surface area and in turn fungus gets food from plant. Therefore, mycorrhizal association is obligatory for Pinus seed germination.
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